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XB-ART-46346
Cell 2012 Sep 28;1511:41-55. doi: 10.1016/j.cell.2012.08.028.
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The transcription factor MEF2 directs developmental visually driven functional and structural metaplasticity.

Chen SX , Cherry A , Tari PK , Podgorski K , Kwong YK , Haas K .


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Natural sensory input shapes both structure and function of developing neurons, but how early experience-driven morphological and physiological plasticity are interrelated remains unclear. Using rapid time-lapse two-photon calcium imaging of network activity and single-neuron growth within the unanesthetized developing brain, we demonstrate that visual stimulation induces coordinated changes to neuronal responses and dendritogenesis. Further, we identify the transcription factor MEF2A/2D as a major regulator of neuronal response to plasticity-inducing stimuli directing both structural and functional changes. Unpatterned sensory stimuli that change plasticity thresholds induce rapid degradation of MEF2A/2D through a classical apoptotic pathway requiring NMDA receptors and caspases-9 and -3/7. Knockdown of MEF2A/2D alone is sufficient to induce a metaplastic shift in threshold of both functional and morphological plasticity. These findings demonstrate how sensory experience acting through altered levels of the transcription factor MEF2 fine-tunes the plasticity thresholds of brain neurons during neural circuit formation.

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Species referenced: Xenopus laevis
Genes referenced: mef2a

References :
Della Santina, A molecular link tethering neuronal responses with the past. 2012, Pubmed